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Molecular studies have instead found evidence for a monophyletic neotropical earless taxon in the basal part of the phylogenetic tree. This, the authors argued, may indicate that Eumantodea arose in Gondwana, or even in South America.

If the structures were obviously strongly damaged in cases where only one or few k,enthal were present, the characters were not encoded for the corresponding taxa. This is very obvious, for example, in many Amorphoscelinae e.

Wieland 2013, The Phylogenetic System of Mantodea

Although our knowledge regarding many taxonomic groups has grown enormously over the last decades, it is still the objective of the Handbook of Zoology to be comprehensive in the sense that text and references together provide a solid basis for further research. Such monogeneric subgroups will be referred to by their generic names throughout the text. Das erste Nymphenstadium der Gottesanbeterinnen besitzt vier Cercusglieder.

Further characters that were mentioned by Roy a: Apomorphic characters for monophyletic taxa within Mantodea are discussed. This will also include those structures that have previously been assumed to have evolved convergently. He named the following diagnostic characters for this group- ing: The fore femora carry an antero-ventral avfs and a postero-ventral pvfs row of spines, and one to four often oblique discoidal spines ds that are missing in Blattodea.


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The head capsule may have various kinds of processes and ridges and carries an aut- apomorphic structure of Mantodea, the so-called scutel-lum or frontal shield. Blepharopsis zoolgisches, antennomere, dorsal view. The praying mantids have retained several plesiomor-phic traits, for instance the presence of three ocelli, the five-segmented tarsi, and the multi-segmented cerci.

Hymenopus coronatus, antennomeres, lateral view.

Mantidae is the largest conglomerate of man-todean species in the typological classification. In Empusi-dae the scutellum carries a more or less strongly developed median ridge see character 8 which, to- gether with the dorsal edge, forms a tapering spine that prominently protrudes the scutellum anteriorly Roy Additionally, the two groups were found in the same larger clade with Amorphoscelinae being the sistergroup of a clade containing several Nanomantinae, Hapaloman-tinae, Tropidomantinae, Liturgusinae, soologisches Paraoxypili-nae.

In the morphological analysis of Neotropical Mantodea by Jantsch The torus intercervi-calis is unique to certain Mantodea and is entirely missing in Blattaria and Isoptera. Furthermore, Exparoxypilus afri-canus was described from Tanzania by Beier This was kkwnthal assumed in the mental analysis published by Grimaldi This condition is also referred to as Antenna imbricata Seifert Further-more, the wing venation has been stated to be most pri-mitive in Chaeteessa Smart In Mantodea, the base of the flagellum may be thick- ened, as for example in Schizocephala bicornis Ehrmann Mantoida maya,lateral view.

Chopard himself suspected that this species might actually belong to Heteronutarsus Chopard All Hymenopodidae except for three genera clustered in a larger clade also containing Phyllotheliinae and Sibyllidae; node that was the sistergroup of Empusidae. A ridge on the clypeus is missing in Blattodea and several Mantodea including Chaeteessa, Mantoida, Metal-lyticus, Amorphoscelidae and Eremiaphilidae, among others. A pointed elongation on each compound eye is exhib- ited by several species in this study [Acanthops Fig.


Similar density of cuticula and amber matrix led to a similar absorption of photons and therefore to a similar brightness of the two substances in the dataset.

Thespidae was found to be monophyletic and to include Oligonicinae as well as Haaniinae, nested among the latter. Thus, their body length measures no more than about per cent of the adult body length. Relationships that are also supported with reservations by morphological data are the basal dichotomy between Chaeteessa and the remaining Mantodea and the monophyly of Artimantodea with reservations regarding the position of Peaktikum.

Pseudocreobo-tra wahlbergii, antennomeres, lateral view.

Die Untersuchung von Nymphen des ersten Stadiums ermglichte fr mehrere Arten neue Interpretationen von Struk-turen bei erwachsenen Tieren. The scutellum of Metallyticus is horizontally folded to an angle of nearly 90 degrees in lateral view Wie-land a: This taxon currently comprises ten genera Roy a: Antennae of female, largest number of antennomeres: As it has been generally assumed that short prothoraces in Mantodea represent the plesiomorphic state, this further contradicts the Mantoidea-hypothesis.

Four cercomeres are present in the first instar nymphs of extant Mantodea.

These characters can also be found in other mantodeans, e. Authors of the taxa studied can be found in Tab. Howe-ver, their findings could not contribute to solve the phy-logenetic relationships of the three subgroups of Amor-phoscelidae.

Volume 1: Evolution, Systematics, and Biogeography

Tropidomantinae as their sister taxon. For those taxa only zoilogisches gender was encoded in the data matrix if no literature data was available. Another important fossil species is Santanmantis axelrodi.